Interstellar yomwe ikubwera njira yamasewera Galactic Civilizations IV ikuwonjezera mabelu atsopano osangalatsa a danga ndi malikhweru a mlengalenga pakubwereza kwake kwachinayi, kuphatikiza The Navigators. Mpikisano watsopano (omwe ukupezeka kuti useweredwe pompano pamayeso a alpha), The Navigators ndi anthu oyendayenda omwe adadzipereka kufunafuna ndikufufuza zinsinsi za Precursors.
Zofuna zawo zakhala zikuchitika kwa nthawi yayitali kotero kuti ayiwala komwe adachokera - mbiri yawo komanso komwe adachokera zidatayika chifukwa cha kuwonongeka kwa nthawi. Nthano imanena za mtundu wawo 'kukhudzidwa' ndi Precursors kale, zomwe zingafotokoze ulendo wawo wautali wofunafuna chidziwitso. Cholowa cha Navigators ndi mitsinje yapansi panthaka, yomwe ndi njira zofunika kwambiri zomwe zimagwirizanitsa magawo atsopano a Galactic Civilizations IV.
A Navigators aganiza zothetsa moyo wawo wosamukasamuka chifukwa apeza zomwe amakhulupirira kuti ndi dziko loyambirira la Precursor: chiwonongeko chophulitsidwa cha pulaneti yotchedwa Arnor. Poyerekeza ndi mwana wina wa Galactic Civilizations IV, a Terrans, Navigators amapereka chimodzi mwazinthu zodziwika bwino zoyambira pamasewerawa ndipo adzawonetsa akale omwe ali ndi vuto latsopano losangalatsa kuposa kale.
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